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I'm not sure when this happened, but this strange button has suddenly appeared. It can be clicked but it doesn't do anything. It's not listed in my actual bookmarks. I've disabled all my extensions and it's still there. I've deleted the contents of my bookmarks toolbar and it still remains.

Anybody seen this before? Any ideas how to get rid of it?

That's the item that determines where your bookmark bar is. Try rightclick on an empty space on your any of your bars and click customize. See if you can remove it like that.

Right click near it or on it. Customise. Drag it off.
Done =)

Hmmm, I can drag it off, but all the bookmarks in the toolbar come with it and I'm left with an empty toolbar. When I drag it back on again the button is still there.

Yup, I'm having this problem, too. Not too sure what's causing it
I'm on the Firefox IRC channel trying to see if anyone else knows what's going on.
If I figure it out, I'll be sure to post back

What's it doing there when the toolbar isn't being customized though?

After spending some time in Safe Mode, then disabling then re-enabling everything, it turned out to be an incompatibility with 3.5b4 and the Strata Reloaded theme. Shame... I really liked that theme.

Edit: Double post.

Btw, v1.7.4 of Strata Reloaded works fine